MD and PhD in Human Genetics from Yale University in 1976 and 1977.

Postdoctoral work on Harvard (Dana-Farber) and Sloan-Kettering Cancer Center, 1977 - 1982

Asst Professor, then Assoc Professor of Pathology at Boston University, 1982 - 1990

Professor of Pathology at University of Michigan, 1990 - present